Plant Systematics

1st chapter

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1- eubacteria2-archaea3-eukaryotes
1- true bacteria, mitochondria, chloroplasts2- crenarchaeota (extremophiles, Sulfolobus), euryarchaeota (halophiles and methanobacteria), related taxa3- protists, plants, fungi, animals
Plants=
One branch in the evolutionary tree of life filled iwth organisms that actively convert CO2 and water into carbohydrates.
---great diversity in kinds of evolutionary status of plants: (a&b?)
--these groups share a number of features:
A. green algae (chlorophytes), charophyean algae ans charophytesb. land plants
--1-chlorophyll a and b2- storage of carbohydrates, usually as starch, within chloroplasts with two membranes3-presence of anterior whiplike flagella, often modified or loss.
Land plants (some live in water) = ?--characterists?
Embryophytes, closest relatives to Charophytes-thick walled spores-embryonic stage in life cycles-specialized structures that protect gametes: 1. archegonium for eggs 2. antheridium for sperm-a cuticle, waxy layer covering the epidermal cells-numerous DNA characteristics that show members of this group are a single branch of the tree of life
Two main groups of land plants:
1- Bryophytes: liverworts: two forms, thalloid and leafy, have stomataMossesHornworts2- Tracheophytes:-largest plant group with 260,000 known spp-members from major vegetation types across earth's terrestrial surface-largest group is angiosperms or flowering plants^---provide lions share of our food and medicine, etc
Systematics
The science of organismal diversity--part of the field of taxonomy
Taxonomy
Classification, description and naming of organisms usually using a hierarchal system.--oldest science= poisonous vs. edible--great deal of disagreement-terms are often used interchangeably.
Systemasists are interested in the evolutionary relationships or kinships between organisms.
---is a study of biodiversity, and its evolutionary history*** plant systematics definition-fundamentally trying to discover the branches of the evolutionary tree of life and document changes that have occurred through time. --systemasists have and continue to develop a system of classification to help us understand Earths diversity and evolutionary/genetic responses
Basic objectives of plant systematics:
1- Provide a convenient method of identifying, naming, and describing plant taxa2- Construct a system of classification that attempts to express phylogenetic relationships and consequently has predictive value.3- Produce an inventory of plant taxa at various scales of interest from local to global=produce floras.4- Develop an understanding of evolutionary processes and relationships.
Stages of plant systematics:
1- Pioneer- discovery and collection phase, still going on in tropics.2- Consolidation- synthesis, mostly based on morphology of field and herbarium specimens in preparation of floras and form morphology/structurally based classification systems.3- Experimental or biosystematic- analysis of wide variety of things such as reproductive systems, variation patterns, chemical, cytological, genetic, fertility relationships or evolutionary patterns.4- Encyclopedic or holotaxanomic- analysis and synthesis of all info and types of data to develop classification systems based on phylogenetic relationships.------stages 1 and 2 are often called alpha taxonomy------stage 4 is often called omega taxonomy.
1-Theory neutral school
(school of systematics)-methodology is theory neutral, they are just describing the similarity and differences between groups of org's around us.---science based on evidence at hand
2- Theory-grounded or based school
(school of systematics)-grouping represented by theorized or hypothesized branches in the evolutionary tree.---based on framework which allows one to hypothesize about possible missing entities we have not observed.---but can predict framework provided by evolutionary tree.
Science of systematics is based on the ability to produce a framework based on the concept of phylogeny:
=the evolutionary history of an organism or group of organisms1-modern molecular techniques and comp tech have revolutionized ability to discern phylogenies.2-example: a group called "Deep Green" was a 5 yr project to construct a comprehensive phylogeny of green plants, by 200 scientists in 12 countries, have used DNA analysis to radically question how green plants are related.
"Deep Green" has overturned several previously held theories:
1- overturned the traditional belief that the "land-plant invasion" was lead by seawater plants. Instead, found that primitive freshwater plants provided the ancestral stock from which all green land plants are descended.2- Also found that Amborella= trichopoda (New Caledonia) one of the oldest living angiosperm, similar to first ancestral angosperm, before it was thought that Magnolias (magnolids) or waterlillies (Nymphales) w/multiple flower parts were the closest relatives to the ancestral angiosperm.
Modern systematics utilize the following:
1- DNA sequencing in genes that readily mutate2- Paleobotany from Miocene3- Advances in comp tech4- biochemistry5- traditional morphology 6- advances in cladistics = characteristics plants have in common